Do dogs have moral intuitions? I'd say yes based on how they respond to failing to reward tricks, giving more treats to dog2, dogs' capacity for self sacrifice, etc.

Do chimps/bonobos have moral intuitions? In addition to the example provided in the podcast, the two primates have very different strategies for conflict resolution despite their genetic proximity, and I have a hard time identifying the point in human evolution that triggered some sort of deontological obligation.

Should we punish lions for eating cubs sired by other lions? Although they are warm-blooded like us, this is normal behavior for the lion. We might say lions don't have the capacity for morality, but that requires distinguishing them from primates, whales, dolphins, rats, etc.

But lets put aside the premise that humans are categorically distinct as moral angents. If we discovered an alien species (perhaps descended from ravens or parrots rather than chimps), could we reasonably judge them by our own standards? What if the conditions were so much harsher than earth that morally appalling behavior was necessarily required for that species to survive? (Relatedly, do we judge other humans who bend/break moral norms in the name of survival, as many survivors of genocide or war have done?).

IMO, moral codes are path dependent, adaptive, and emergent. We see this biologically with the examples provided in the podcast, but also in human history and political philosophy.

We can even take this principle one further into the realm of political philosophy. Suppose a sovereign has a (deontologicsl) moral code that prohibits taking violent action against a rebellious faction that is ready and willing to commit violence. Should the sovereign adhere to their code even where the rebels will impose a moral order that would be horrifying to the sovereign? I don't find this example much different than an animal with poor self-preservation instincts, but some of my comments below go into more details about my premises and corrolary arguments etc.